To extend the actuarial analogy, individuals can be thought of as life-insurance underwriters. An individual can be expected to invest or risk a certain proportion of his own assets in the life of another individual. He takes into account his relatedness to the other individual, and also whether the individual is a 'good risk' in terms of his life expectancy compared with the insurer's own. Strictly we should say 'reproduction expectancy' rather than 'life expectancy', or to be even more strict, 'general capacity to benefit own genes in the future expectancy'. Then in order for altruistic behavior to evolve, the net risk to the altruist must be less than the net benefit to the recipient multiplied by the relatedness.

Richard Dawkins -The Selfish Gene.


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